从蓼科植物的孢粉类型角度,研究了中国蓼科花粉类型的地理分布格局及其与生态因子的关系。根据中国蓼科植物赖以生存的生态因子,得出中国蓼科花粉类型分布区的主要生态因子,包括地理位置(分布中心)、海拔高度、年降水量、年积温及生境数量。在此基础上,根据同一区域内相同或相似的生态环境条件下分布的现代蓼科各种花粉类型,确定一定花粉类型组合所指示的现代气候和环境,为利用地层中蓼科化石花粉重建古气候、古环境及气候变迁提供了现代孢粉学证据。
In the present paper, geographic distribution patterns of pollen types of Polygonaceae and their relationship to ecological factors are first introduced. Geographic ditribution centre of Atraphaxis-type,Calligonum-type and Aviculare-type is located in the northwestern China, but they differ in other factors. Atraphaxis-type and Calligonum-type are in the habitats of gobi desert and sandy land with altitudes from 300 to 800 m, annual precipitation from 20 to 200 mm, annual accumulated temperature from 1600℃ to 3400℃;Aviculare-type is in the habitats of arid hillside,calcareous desert and flood grassland with altitudes from 400 to 2000 m,annual precipitation from 200 to 800 mm,annual accumulated temperature from 1600℃ to 4500℃.Geographic ditribution centre of Rheum-type, Koenigia-type and Acetosa-type is located in Northwest China and Southeast China, but they differ in other factors. Rheum-type is in the habitat of mountain slope of cold high mountain area with altitudes from 2000 to 4000 m, annual precipitation from 50 to 400 mm, annual accumulated temperature from 1600℃ to 2000℃; Koenigia-type is in habitat of meadow in summit of a mountain with altitudes from 2000 to 4900 m, annual precipitation from 300 to 800 mm, annual accumulated temperature from 600℃ to 3000℃;Acetosa-type is in habitats of wetlant in mountain valley and near river with altitudes from 450 to 3100 m, annual precipitation from 200 to 800 mm, annual accumulated temperature from 1600℃ to 6000℃.Geographic ditribution centre of Aconogonon-type, Cephalophilon-type and Polystachyum-type is located in Southwest China. They are in habitats of shrubbery and grassland on mountain slope and wetland in mountain valley with altitudes from 1000 to 3000 m, annual precipitation from 400 to 1600 mm. Geographic ditribution center, habitats,altitudes and annual precipitation of Fagopyrum-type resemble that of Aconogonon-type,Cephalophilon-type and Polystachyum-type,but they differ in annual accumulated temperature. Fagopyrum-type is from 4500℃ to 8000℃; the others from 1600℃ to 8000℃.Geographic ditribution centre of Bistorta-type and Aubertii-type is located in Southwest China and Central China, but they differ in other factors. Bistorta-type is in habitats of meadow in summit of a mountain and wetland in mountain valley with altitudes from 1300 to 5000 m, annual precipitation from 400 to 1200 mm, annual accumulated temperature from 1600℃ to 8000℃; Aubertii-type is in habitats of grassland on mountain slope, shrubbery in mountain valley with altitudes from 900 to 3200 m, annual precipitation from 400 to 800 mm, annual accumulated temperature from 1600℃ to 5000℃.Geographic ditribution centre of Pteroxygonum-type is located in Qinling in habitats of crevice in rock and shrubbery on mountain slope with altitudes from 600 to 2000 m,annual precipitation from 400 to 800 mm,annual accumulated temperature from 3400℃ to 4500℃.Geographic ditribution centre of Persicaria-type and Tovara-type is located in East China and Central China in habitats of wetland near gully with altitudes from 30 to 2500 m, annual precipitation from 400 to 1600 mm,annual accumulated temperature from 4500℃ to 8000℃.Geographic ditribution centre of Amphibium-type and Convolvulus-type is located in Southwest China, Central China and Northeast China, but they differ in other factors. Amphibium-type is in habitats of shallow water in margin of lake and near gully with altitudes from 50 to 3700 m, annual precipitation from 800 to 1600 mm, annual accumulated temperature from 1600℃ to 6000℃;Convolvulus-type in habitats of grassland on mountain slope and wetland in mountain valley with altitudes from 100 to 2800 m, annual precipitation from 300 to 1600 mm, annual accumulated temperature from 1600℃ to 6000℃.Geographic ditribution centre of Tiniaria-type is located in Southwest. China, Central China and East China in habitats of shrubbery in mountain valley with altitudes from 200 to 3000 m,annual precipitation from 800 to 1600 mm, annual accumulated temperature from 4000℃ to 8000℃.Distributions of the pollen types indicate arid and desert climate in Northwest China with assemblage of Atraphaxis-type and Calligonum-type;warm, humid and subtropical climate in Southwest China with assemblage of Aconogonon-type, Cephalophilon-type,Polystachyum-type, Acetosa-type and Fagopyrum-type; high and cold climate in Qinghai-Tibet Plateau with assemblage of Koenigia-type and Rheum-type; warm humid subtropical climate and tropical monsoon climate in southern coastal area with assemblage of Persicaria-type and Tovara-type; warm and humid climate in Central China and East China with assemblage of Persicaria-type, Bistorta-type, Tiniara-type and Tovara-type; warm and semi-humid climate in North. China with assemblage of Bistorta-type, Aubertii-type and Pteroxygonum-type; humid, semi-humid, middle warm and monsoon climate in Northeast China with assemblage of Amphibium-type, Convolvulus-type and Sibiricum-type.
[1] 包伯坚,李安仁.中国木蓼属的研究兼论木蓼族的系统[J].植物分类学报, 1993,(2):127~139.
[2] 周忠泽,许仁鑫,庄永龙,等.蓼科花粉外壁超微结构的研究[J].植物分类学报,2000,38(5):446~451.
[3] 周忠泽,张小平. 孢粉地理学的研究方法[J].地理科学, 2000,20(2): 172~175.
[4] 周忠泽,鲁润龙,郑艳燕.中国蓼科花粉萌发孔的平行进化[J].中国科学技术大学学报, 1999,29(5):569~577.
[5] 周忠泽,江光怀,鲁润龙.中国蓼科孢粉地理学的研究[J].中国科学技术大学学报,1999,29(4): 464~470.
[6] 周忠泽,张小平,鲁润龙,等.中国蓼科一新的花粉类型的发现及其演化地位和分类上的意义[J].中国科学技术大学学报, 1999,29(2): 195~198.
[7] 周忠泽, 鲁润龙, 葛磊,等.安徽蓼科花粉形态的研究[J].植物研究, 1999, 19(1):17~33.
[8] 张小平, 周忠泽.中国蓼科花粉的系统演化[M].合肥: 中国科学技术大学出版社,1998.
[9] 王伏雄,钱南芬,张玉龙,等. 中国植物花粉形态[M].北京:科学出版社, 1999. 318~324.
[10] 王建新,冯志坚.中国蓼属植物花粉形态的研究[J].植物分类学报, 1994, 32(3): 219~231.
[11] Hong S-P,Hedberg O. Parallel evolution of aperture numbers and arrangement in the genera Koenigia, Persicaria and Aconogonon (Polygonaceae) [J]. Grana, 1999, 29: 177-184.
[12] 刘会平,唐晓春,王开发,等.神农架北坡表土常见花粉的R值研究[J].地理科学,2001, 21(4):378~380.
[13] 许清海,杨振京,崔之久,等.赤峰地区孢粉分析与先人生活环境初探[J].地理科学,2002, 22(3): 324~329.
[14] 张玉兰,杨永兴.中全新世以来黑龙江同江地区的孢粉组合与植被气候演化[J].地理科学,2002,22(4):426~429.
[15] 杨永兴,王世岩.8.0?ka B.P.以来三江平原北部沼泽发育和古环境演变研究[J].地理科学,2003, 23(1): 32~38.
[16] 国家自然地图集编纂委员会. 中华人民共和国国家自然地图集[M]. 北京:中国地图出版社, 1999.
[17] 吴征镒(主编).中国植被[M]. 北京:科学出版社, 1980.
[18] 李安仁. 中国植物志,25(1)——蓼科[M]. 北京:科学出版社,1998.
[19] Godwin H. The history of the British Flora. A factual basis for phytogeography[M]. Cambridge: Cambridge Uiv Press, 1975. 2 nd ed.541.
